Sys.InvalidOperationException: Sys.InvalidOperationException: Could not find UpdatePanel with ID ''. If it is being updated dynamically then it must be inside another UpdatePanel.

I have converted asp.net3.5 project to .net4.5.1 (VS2013) but i get an error like "Sys.InvalidOperationException: Sys.InvalidOperationException: Could not find UpdatePanel with ID ''. If it is being updated dynamically then it must be inside another UpdatePanel.". its only get when running through VS2013. I  didn't get this is issue  when publishing the application.